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    One of the 3 sequels of "An American Tail", this takes place before "An American Tail: Fievel Goes West". Fievel Mouskewitz and his family, Tanya, his sister, Mama, Papa, and baby sister, Yasha, are trying to get used to life in New York City in America. But it's not long before Fievel, his friend Tony Toponi, a scientist named Dithering, and his assistant, Scuttlebutt, find out that a map found in an old subway, leads to "...our greatest treasure." They go on an expedition to find the treasure, and after facing booby traps, they come across indians living underneath New York City. Fievel and Tony meet the indian cheif's daughter, Cholena, and she shows Fievel that their greatest treasure is their story told in symbols. The cheif sends her to the surface to see if the mice up there have changed their ways and haven't brought disease and cats like in the old days. Unfortunately, Scuttlebutt works for the villians of the movie, the owners of a cheese factory, Mr. O'bloat, Mr. Toplofty, and Mr. Grasping. The villianous trio, keeping an eye on Fievel's papa, corrupt the mice of Fievel's community and become an angry mob and look for Cholena. Fievel, Tony, and Tanya try to get Cholena back to her tribe under the ground, but realize that they are being pursued by police cheif McBursque, Scuttlebutt, and the other police officers. Realizing this, they must be ready. There's no turning back now...